Key Laboratory of Microbial Technology for Industrial Pollution Control of Zhejiang, Province, College of Environment, Zhejiang University of Technology, Hangzhou, 310032, China. Zhejiang University of Water Resource and Electric Power, Hangzhou, 310018, China. Key Laboratory of Microbial Technology for Industrial Pollution Control of Zhejiang, Province, College of Environment, Zhejiang University of Technology, Hangzhou, 310032, China. Electronic address: shihanzhang@zjut.edu.cn.